nodosa present in Mar Menor at least
in the last decades. Caulerpa contains
high levels of toxigenic secondary metabolites and contributes loads of labile
organic matter to the sediments. Decomposition of this organic matter fuels
anoxic processes and increased levels of
reduced carbon, nitrogen and sulphur
compounds.
Fig. 1. Maps of macrophytes assemblages in Mar Menor lagoon before and after the 2016 anoxic
event. Drastic changes in the distribution of the two dominant macrophyte species, the green alga
Caulerpa prolifera and seagrass Cymodocea nodosa, are illustrated
thresholds, known in ecological theory
as tipping points, had been crossed.
Tipping points are levels of pressure
beyond which the ecosystem can no
longer cope with change, and suddenly
shifts from one state to another (ecosystem shift), usually more deteriorated,
but equally stable state [5].
How did this iconic ecosystem
reach this condition?
In the first place, it is necessary to know
the environmental framework of Mar
Menor, a shallow (maximum depth 7m)
hypersaline coastal lagoon with a catchment basin of 1,300 km2 and a water
renewal time of about 1 year [6]. According to available information on the
semi-arid climate prevalent in the SE
Iberian Peninsula, the only freshwater
inputs to the Mar Menor are from runoff
associated with a few torrential rains
which occur every year (250 mm year1), and some more diffusive groundwater inputs. Recently developed models
[7] indicate that the current input from
runoff is quantitatively more important
than that from aquifers. Nevertheless,
the little and dispersed data available
are not sufficient to provide reliable
data for lagoon management.
2
1. Impact of the Estacio channel
widening
In 1973, widening of the Estacio Channel, one of the five natural connections
(golas) between the lagoon and the
Mediterranean, was carried out [8].
This public work, now considered a
landmark of the anthropogenic transformation of the lagoon, changed the
hydrodynamic regimen and led to a decline in salinity, from 50-52.5 in the early 1970s to the current values (42-48)
which show an interannual variability
determined by the frequency and intensity of the DANAS or gota fría, i.e.
intense cold spells bringing heavy rainfall. The increased exchange favoured
the entry of some Mediterranean species and the decline of some others,
members of the autochthonous fauna
and flora. Water exchanges through the
Estacio channel are now the main control of the lagoon hydrodynamics. One
of the most favoured introduced species
was the green macroalga Caulerpa prolifera, a well-known opportunistic macrophyte able to take up nutrients very
efficiently and colonize extensive areas
rapidly. This species has properties conferring the ability to outcompete native
macrophyte assemblages, in particular
meadows of the seagrass Cymodocea
2. Introduced macroalgal species and
uncontrolled wastewater discharges
There is no argument about the increased primary production and deposits of organic matter sedimented in the
lagoon during the last 60 years [9], but
the negative impact of these changes
on the ecosystem may have been overestimated, and conclusions have not
been based on scientific evidence. A
good example of this criticism concerns
the supposed regression of native seagrass meadows, in particular C. nodosa,
as a consequence of the expansion of
C. prolifera [10]. Recent studies have
confirmed that angiosperms did not
decline during the decades preceding
the collapse. Conversely, these studies
provided evidence of the coexistence of
the two macrophytes at least during the
last forty years before the 2016 event
[11] (Fig. 2) indicating a benthic primary production enhancement. Increased
primary production supported by high
availability of nutrients is an intrinsic
feature of eutrophic ecosystems [12].
Therefore, man-made changes induced
on the lagoon hydrodynamics were not
the only factor that explained the expansion of C. prolifera.
Uncontrolled discharges of untreated wastewaters from urban and tourist
developments took place years before
the channel widening. In the 1990s,
the wastewater treatment plan for Mar
Menor was completed and wastewaters
were no longer flushed into the lagoon
(at the expense of being discharged into
adjacent Mediterranean waters). Nevertheless, problems related with high
nutrient inputs persisted or even increased due to the rapid development
of intensive agriculture which started in
the 1950s [13]. This kind of agriculture
replaced the traditional drylands at the
expense of aquifer overexploitation.
The new agricultural practices marked
another step towards deterioration of
the lagoon [14]. Today, the irrigation
area occupies 40% of the total drainHARMFUL ALGAE NEWS NO. 70 / 20221
Harmful Algae News An IOC Newsletter on Toxic Algae and Algal Blooms No. 70 - July 2022 https://hab.ioc-unesco.org/ Mar Menor lagoon: an iconic case of ecosystem collapse Content Featured articles Mar Menor lagoon: an iconic case of ecosystem collapse, Juan M Ruiz, Jaime BernardeauEsteller, M Dolo
nodosa present in Mar Menor at least in the last decades. Caulerpa contains high levels of toxigenic secondary metabolites and contributes loads of labile organic matter to the sediments. Decomposition of this organic matter fuels anoxic processes and increased levels of reduced carbon, nitrogen and
age basin and is a major source of European winter vegetable production. But the transferred water resources, clearly insufficient to sustain such production, had to be complemented with aquifers that had suffered previous overexploitation and became brackish. These brackish aquifers needed treating
Fig. 3. Satellite image (Sentinel 2) after torrential rainfall in September 12th and 13th in the Mar Menor watershed. Tons of terrigenous sediments, carbon, nitrogen and phosphorous are dragged by water runoff from agricultural lands into the Mar Menor lagoon (downloaded from https://www. copernicus
provided by President and Staff of the harbours Club Nautico Lo Pagán, Club Náutico La Puntica and Centro de Actividades Náuticas (San Pedro del Pinatar, Murcia, Spain). References 1. Ruiz JM et al 2020. Informe de asesoramiento técnico del IEO, 165pp 2. Belando MB et al 2019. Front. Mar. Sci. Conf
The Mar Menor Oyster Initiative, a strategy to prevent algal blooms in a eutrophic lagoon in Spain Fig. 1. Map of study in Mar Menor, Murcia, SE Spain Marine ecosystems are exposed to a wide range of pressures including water quality degradation, habitat decline, overfishing and climate change, in
flats, located in the south basin of the lagoon. The rationale of this project is to involve all interested stakeholders for successful large-scale restoration programs, which need public and political support, research, and outreach actions [13]. Acknowledgements Project RemediOS is developed with
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