ever the information presented in publications is not always easy to interpret.
Toxic blooms often contain several species and studies on individual species
are therefore required to determine the
species responsible for the toxin production. In addition, species are sometimes difficult to identify and the species
concept in cyanobacteria is a challenge.
Nomenclatural Problems
There are four specific problems facing
the Task Team:
Problem 1. The Alexandrium
problem
Alexandrium is one of the commonest used generic names for toxin-producing HAB dinoflagellate species, but
there was initial disagreement about
the name to be used for the tamarense
group and its allies: Alexandrium Halim,
Protogonyaulax F.J.R. Taylor, Gessnerium
Halim or Gonyaulax Diesing. An important step was made towards avoiding
further confusion at the HAB Conference in Lund, Sweden, in 1988 when the
taxonomic expert, Prof. Enrique Balech
from Argentina, was invited to discuss
and give his opinion on the taxonomy
of the group. After a detailed presentation, Balech concluded that all species
should be included in the genus Alexandrium Halim. This resulted in heated
discussion from/in the audience but
objections raised were rejected by Prof.
Balech. The chairman of the session,
Prof. Karen Steidinger, finally concluded
the discussion by asking all persons in
the audience to agree on using only Alexandrium in the future. This proposal
was adopted and the name Alexandrium
has subsequently been used by scientists working on HABs.
However in 2019 it was discovered
that a species of the poorly known genus Centrodinium nested within the
genus Alexandrium in the molecular
trees provided by Li et al [1]. Species
belonging to both genera formed a
monophyletic group. As Centrodinium
was described by Kofoid in 1907 [2] but
2
Alexandrium by Halim only in 1960, the
old name Centrodinium takes priority if
the two genera are to be merged.
As changes of well-known names
are a source of confusion, the botanical code establishes that well-known
names of particular importance may be
conserved: Conservation aims at retention of those names that best serve stability of nomenclature (International
Code of Nomenclature for Algae, Fungi
and Plants 2018, Art. 14, 2). The case
of Alexandrium versus Centrodinium is
such a case. A formal proposal to conserve the name Alexandrium instead of
Centrodinium is now being prepared
to be submitted to the General Committee of the International Code of Nomenclature for Algae, Fungi and Plants.
Subsequently splitting of Alexandrium
was suggested [3]. An article containing arguments against this move has
been submitted by Mertens et al. (pers.
comm.). Until the problems have been
settled, the IOC List of Harmful Algae
will continue to use the name Alexandrium in the context of the 1988 decision.
Problem 2. The Cochlodinium
polykrikoides problem
One of the most harmful fish killers
has been referred to as Cochlodinium
polykrikoides since its description from
Puerto Rico in 1961. Huge fish kills
have occurred in East Asia as a result of
blooms of this species and South Korea
has been particularly badly impacted.
The species name was recently changed
to Margalefidinium polykrikoides (Margalef) F. Gómez, Richlen & Anderson
2017 [4]. To change the name of wellknown species always causes confusion
and misunderstandings, and the problem in this case is as follows. A genus
is nomenclaturally anchored to its type
species and the type species of Cochlodinium is Cochlodinium strangulatum
(F. Schütt) F. Schütt 1896. Like many
other unarmoured (naked) dinoflagellates, Cochlodinium is and has always
been a heterogenous genus. It comprises both chloroplast-containing and
chloroplast-lacking species, and molecular data have shown, not surprisingly,
that the chloroplast-containing species
Cochlodinium polykrikoides Margalef
is not closely related to the heterotrophic, chloroplast-lacking type species
C. strangulatum. They should therefore
not be retained in the same genus, and
the transfer of C. polykrikoides to the
new genus Margalefidinium is a logical
proposal. Due to the economic importance of this species, conservation of
the name Cochlodinium polykrikoides
would cause much less confusion. Conservation can be attained by formally
proposing to the General Committee of
the International Code of Nomenclature for Algae, Fungi and Plants that the
genus Cochlodinium be conserved with
C. polykrikoides as type species, to replace C. strangulatum. A proposal to
that effect is now being prepared and,
if accepted, will conserve the name Cochlodinium polykrikoides. The species C.
strangulatum will then need to be given
a new generic name but given the fact
that it is a little-known species, a new
generic name for this species will cause
little confusion. A species from the Far
East related to Cochlodinium polykrikoides is Cochlodinium catenatum Okamura described from Tokyo Bay [5]. If the
proposal to conserve the genus Cochlodinium with Cochlodinium polykrikoides
as type is accepted, C. catenatum will
again be the correct name of that species.
Problem 3. Prorocentrum rhathymum and P. mexicanum
Gómez et al [4] have recently claimed
that the species Prorocentrum rhathymum Osorio-Tafall and P. mexicanum
A.R. Loeblich, Sherley & R.J. Schmidt
are synonyms. The question of possible
synonymy goes back to Steidinger in
1983 [6] who claimed, four years after
P. mexicanum was described by Loeblich et al [7], that the two taxa were
synonyms. Faust [8] went on to state,
incorrectly, that Loeblich et al, when describing the new species P. rhathymum,
HARMFUL ALGAE NEWS NO. 64 / 2020
Harmful Algae News An IOC Newsletter on Toxic Algae and Algal Blooms No. 64 - June 2020 www.ioc-unesco.org/hab The IOC Taxonomic Reference List of Harmful Microalgae The creation of the IOC Taxonomic Reference List of Harmful Microalgae was first discussed 23 years ago at the Fourth Session of the
ever the information presented in publications is not always easy to interpret. Toxic blooms often contain several species and studies on individual species are therefore required to determine the species responsible for the toxin production. In addition, species are sometimes difficult to identify
considered this to be identical to the previously described P. mexicanum [9]. This statement makes little sense, and Loeblich et al in fact do not mention P. mexicanum in their article. Faust [8] illustrated material which resembled P. rhathymum using the radical arrangement of some of the trichocys
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Fish kill in numbers [2] 13 000 ton fish 7.5 mill salmon 80 mill EUR 14 companies Fig. 2. Map of Norway showing the area where the Chrysochromulina leadbeateri bloom occurred causing massive fish kills in May-June 2019 References 1. Fiskeridirektoratet 2019. https:// www.fiskeridir.no/Akvakult
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Fig. 2. Maximum likelihood (ML) molecular phylogenetic tree of Heterocapsa minima and H. steinii strains isolated from coastal areas of New Zealand based on the LSU rDNA D1/ D2 sequences (20 sequences, 700 positions). The alignment site corresponded to the 69768 bp site of a sequence from H. steinii
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Table 1. Physico-chemical parameters of P. quadridentatum bloom area in Cochin Estuary Parameters Water Temperature (oC) Salinity (psu) Dissolved Oxygen (ml L-1) Nitrate (μmol L-1) Silicate (μmol L-1) Phosphate (μmol L-1) Chlorophyll a (mg m-3) 28 20 2.01 10.6 23.3 1.3 27.5 higher turbidity, lowe
Are mesophotic seamounts reservoirs for potentially toxic dinoflagellates associated with Ciguatera poisoning? A case study from the SW Indian ocean (expédition La Pérouse, 2019) Fig. 1. Location map of La Pérouse seamount off La Réunion. In the western Indian Ocean, a Ciguatera Fish Poisoning (CF
Using Machine Learning to Observe Abundance Patterns of the Dino flagellate Noctiluca scintillans in the Western English Channel Noctiluca scintillans is a Harmful Algal Bloom (HAB) species with a wide geographic distribution. It frequently blooms and causes negative impacts on marine ecosystems [1]
References 1. Sarma VVSS et al 2019. Mar Pollut Bull 138: 428-436 2. Faust MA & RA Gulledge 2002. Contributions for the US National Herbarium 42: 1-144 3. Graham MD et al 2018. Limnol Oceanogr-Meth 16: 669-679 Author Christian Bamber, The Marine Biological Association, Citadel Hill, Plymouth PL1 2PB
International validation and recognition of method for paralytic shellfish toxins in bivalve molluscs Food safety scientists from Cefas (UK) and Cawthron Institute (New Zealand) have led an international study over the past four years to gain international recognition for a new method to quantify pa
Aotearoa/New Zealands nationally significant Cawthron Institute Culture Collection of Microalgae (CICCM) The CICCM is critical to international research projects and the 500 isolates of microalgae and cyanobacteria in the collection have been sourced from oceans, lakes and rivers in New Zealand, the
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Typical benthic habitat with macroalgae and limestone rubble observed at ~ 60m depth on La Pérouse seamount, Indian Ocean (Photo courtesy of L. Ballesta). Eds-in-chief Beatriz Reguera, IEO, Vigo, Spain Eilen Bresnan, Marine Scotland, UK Regional Editors Caribbean: Ernesto Mancera jemancerap@unal.e