Fig. 2. Maximum likelihood (ML) molecular
phylogenetic tree of Heterocapsa minima and
H. steinii strains isolated from coastal areas
of New Zealand based on the LSU rDNA D1/
D2 sequences (20 sequences, 700 positions).
The alignment site corresponded to the
69768 bp site of a sequence from H. steinii
strain UTKG7 (clone 1, MF423356). The ML
and neighbour joining (NJ) phylogenies were
analysed using the best-fit models, GTR+G+I
and TN93+G models, respectivelywith bootstrap analyses performed with 1,000 replicates.
Sequences obtained in the present study
are shown in blue and red font. Sequences
corresponding to the type material of each
species are shown in bold font. Nodal support
represents ML/NJ bootstrap values. Nodal
supports that were under 50 are shown as
hyphens. Nodes that were not present in the
NJ tree are labelled as np.
the temperate North Atlantic. However,
details of morphological and molecular
characterisations were not reported
[11]. In 2009, H. minima strain JK2 was
established from Bantry Bay, Southwest
Ireland in the temperate North Atlantic,
potentially representing the same water
mass as the type locality of H. minima
[10], and identified by a combination of
morphological and molecular phylogenetic analyses [12]. Recently, in 2016, a
H. minima strain HMMJ1604, was established and identified by molecular and
morphological analyses from Mijo Port,
South Sea of Korea in the temperate
North Pacific. This finding was the first
report of this species from the Pacific
[13]. Therefore, the present study is the
second record of H. minima from the Pacific region and suggests that this species is widely distributed in the temperate zone of the Atlantic and Pacific. To
the best of our knowledge, the present
study is the first record of this species
from the Southern Hemisphere.
In previous studies reporting the occurrence and abundance of H. minima
cells [10-11,13], it was stated that this
species occurred throughout the year
HARMFUL ALGAE NEWS NO. 64 / 2020
and was widely distributed in the Celtic
Sea and Korean waters [10,13]. Because
the cell morphologies of Heterocapsa
spp. are superficially similar in size,
shape and swimming mode to other
small dinoflagellates (e.g., Azadinium
and Biecheleria) under LM observation,
it is difficult to differentiate them and
high magnification microscopy or molecular tools are required [14,7]. Therefore, the determination of accurate cell
concentrations of these taxa from routine samples used for phytoplankton
monitoring is difficult. For example,
the New Zealand Marine Phytoplankton Monitoring Programme reports
these cell types as cf. Azadinium spp.
(= Azadinium-like species) to provide a
conservative report of potentially toxic
species [9]. In the present study, the
H. minima strain CAWD302 was isolated from a seawater sample collected
at Site 4 on 12 December 2018, from
which a cell density of 6,000 cells/L of
cf. Azadinium spp. was reported. In addition to H. minima, two clonal strains
of Biecheleria spp. strains MB11 and
CAWD304 (GenBank accession numbers: LC542926 and LC542925, respectively) were also isolated from the
samples collected at Site 4 on 5 and 12,
December 2018, respectively. To resolve
this issue for H. minima, Lee et al. [13]
developed a species-specific quantitative PCR (qPCR) method to detect and
quantify cells from the Korean seawater samples. To reveal the accurate distribution of H. minima, applying rDNA
sequencing and/or qPCR to isolated
strains or seawater samples collected
from the other climate zones (e.g.,
tropical, subtropical, and/or subboreal
zones), as well as other temperate areas
is required. Additionally, toxin and/or
toxicity assessments are also needed to
further elucidate the potential impacts
of this species.
Acknowledgements
We are grateful to the Ministry of Primary Industries for allowing the use of
samples for research purposes, and to
Tony Bui and Catherine Moisan (Cawthron Institute Natural Toxins laboratory) for phytoplankton notifications
and access to samples. Funding for
this research was from the Ministry for
Business, Innovation and Employment
(Seafood Safety programme, Contract
No. CAWX1801; National Databases
contract No. CAWX0902). M. Balci was
supported by the Scientific and Technological Research Council of Turkey
under a TUBITAK-BIDEB-2219-International Postdoctoral Research Fellowship (Grant no: 1059B191800400) during the present study at the Cawthron
Institute. T. Nishimura was supported
by the Japan Society for the Promotion
of Science Overseas Research Fellowship.
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Authors
Muharrem Balci, Istanbul University, Faculty
of Science, Department of Biology, 34134
Vezneciler, Istanbul, Turkey.
Tomohiro Nishimura, Kirsty F. Smith, Lesley
L. Rhodes, J. Sam Murray & A. Lincoln MacKenzie, Cawthron Institute, 98 Halifax Street
East, Nelson 7012, New Zealand.
Co-first authors contributed equally to this
work.
Emails corresponding authors:
muharrem.balci@istanbul.edu.tr
tomohiro.nishimura@cawthron.org.nz
9
Harmful Algae News An IOC Newsletter on Toxic Algae and Algal Blooms No. 64 - June 2020 www.ioc-unesco.org/hab The IOC Taxonomic Reference List of Harmful Microalgae The creation of the IOC Taxonomic Reference List of Harmful Microalgae was first discussed 23 years ago at the Fourth Session of the
ever the information presented in publications is not always easy to interpret. Toxic blooms often contain several species and studies on individual species are therefore required to determine the species responsible for the toxin production. In addition, species are sometimes difficult to identify
considered this to be identical to the previously described P. mexicanum [9]. This statement makes little sense, and Loeblich et al in fact do not mention P. mexicanum in their article. Faust [8] illustrated material which resembled P. rhathymum using the radical arrangement of some of the trichocys
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Fig. 2. Maximum likelihood (ML) molecular phylogenetic tree of Heterocapsa minima and H. steinii strains isolated from coastal areas of New Zealand based on the LSU rDNA D1/ D2 sequences (20 sequences, 700 positions). The alignment site corresponded to the 69768 bp site of a sequence from H. steinii
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