Bloom of a red tide species Akashiwo
sanguinea in Semerak Lagoon, Kelantan, Malaysia
March 2016: i.e. Pseudo-nitzschia spp.
(potentially toxic species), Chaetoceros,
Skeletonema, and Blixaea quinquecornis
(red tides, fish kills) [3].
In this survey, we confirmed the
blooming species as the dinoflagellate
A. sanguinea belonging to ribotype B.
The potential occurrence and distribution of other ribotypes require further
investigation, and the drivers of the
recurrence of this species remain unknown. Therefore, routine monitoring
along the lagoon is needed for early
warning to minimize losses due to fish
kill events.
Acknowledgements
This work was supported by the Ministry of Education Malaysia HICoE fund
[IOES-2014C] to PT Lim. SN Tan is supported by UMT post-doctoral fellowship.
Fig. 1. Chart of sampling sites at the Semerak Lagoon and cell densities of Akashiwo sanguinea
The species Akashiwo sanguinea is a cosmopolitan naked dinoflagellate known
to form harmful algal blooms (HABs).
This bloom-forming species has been
reported to cause seabird mortality due
to saponification in Monterey Bay, California, USA [1]. In Malaysia, there are no
reports of fish kill incidents caused by
A. sanguinea to date, but the species is
commonly found in Malaysian waters,
especially in aquaculture areas [2-3].
The Semerak Lagoon, located in the
north east of Peninsular Malaysia, is an
important aquaculture area for finfish
and shrimp. On May 23th, 2019, water
discoloration was observed in the lagoon (Fig. 1). Following the incident,
morphological and molecular analyses
were performed to identify the bloom
species. Water samples from nine selected sampling sites (S1S9; Fig. 1)
were collected at 1 m depth using an 8-L
Van Dorn water sampler. The sampling
stations S1 to S3 were located inside, S4
to S8 outside the aquaculture area, and
S9 at the river mouth. Clonal cultures
were established through single-cell
isolation from field samples. Field and
cultured cells were observed using light
microscopy and confirmed the species
as Akashiwo sanguinea. Cells were 60.6
to 81.0 μm long and 46.6 to 60.6 μm
wide with numerous ribbon-like chloroplasts in the periphery (Fig. 2).
Gene amplification of the large subunit (LSU) region and the internal transcribed spacer (ITS) region was perHARMFUL ALGAE NEWS NO. 65 / 2020
formed by polymerase chain reaction.
The species was previously reported
to comprise four ribotypes (A, B, C, and
D) [4]. Our results based on both LSU
rDNA and ITS sequence analysis clearly
indicated that the A. sanguinea population in Semerak Lagoon belonged to
ribotype B; sharing the same ribotype
with the strains found in Tumpat, Kelantan [4]. The LSU phylogenetic inference
revealed that the strains from Semerak
Lagoon shared identical sequences with
those from China, Mexico, Singapore,
and South Korea (Fig. 3A; MP/ML/BI:
99/99/1.0); while using ITS phylogeny,
all strains from Semerak Lagoon shared
identical sequences with one strain
(GSXM02) from Xiamen Harbour, China
(Fig. 3B; MP/ML/BI: 100/100/0.99).
During the bloom event, the highest
density of A. sanguinea was found at S7
(248,658 cells L-1; outside the aquaculture area), followed by S3 (177,773 cells
L-1; inside the aquaculture area), and S8
(164,269 cells L-1; outside aquaculture
area) (Fig. 1); the densities were higher
as compared to the previous occurrences in 2015 and 2016 (highest density of
3,460 cells L-1; [3]). The phytoplankton
community assemblage in the lagoon
was diverse, with a dynamic shifting of
phytoplankton community composition
over time related to the nutrient dynamics in the lagoon. Aside from blooms
of A. sanguinea, several different algal
blooms were encountered between the
study period of September 2015 and
References
1. Jessup DA et al 2009. Plos One 4: e4550
2. Mohd Razali R et al 2015. Malaysian J
Sci 34: 2436
3. Er HH et al 2018. Environ Sci Pollut Res
Int 25: 2294422962
4. Luo Z et al 2017. Harmful Algae 66:
8896
Authors
Suh Nih Tan, Mohd Fadzil Mohd Akhir &
Zainudin Bachok, Institute of Oceanography and Environment, University Malaysia
Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia
Wendy Wee, Chui Pin Leaw & Po Teen Lim,
Bachok Marine Research Station, Institute
of Ocean and Earth Sciences, University of
Malaya, 16310 Bachok, Kelantan, Malaysia
Email corresponding author:
suhnih@umt.edu.my
Fig. 2. Light micrograph of Akashiwo sanguinea from Semerak Lagoon. Scale, 20 μm
15
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